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Type Mutant Strain; Spontaneous Mutation; Species laboratory mouse Generation N27 Appearance
black, ataxic
Related Genotype: a/a Cacnb4lh/Cacnb4lh
agouti, ataxic
Related Genotype: A/? Cacnb4lh/Cacnb4lh
black, unaffected
Related Genotype: a/a Cacnb4lh/+ or a/a +/?
agouti, unaffected
Related Genotype: A/? Cacnb4lh/+ or A/? +/?Description
Mice homozygous for the lethargic spontaneous mutation (Cacnb4lh) are first recognizable at 15 days by their lethargic behavior with gait instability and occasional seizures. The seizures resemble human petit mal seizures. No pathological changes were found in the CNS or in skeletal muscles, but peripheral motor nerves show reduced conduction velocity and prolonged distal latency. There is early thymic involution at 3 to 4 weeks in Cacnb4lh homozygotes, accompanied by decreased lymphocyte count, decreased cell-mediated immunity, and increased levels of serum IgG1. The defects in the immune system tend to disappear by 2 months of age in mice that survive. In addition to neurological and immunological defects, homozygous lethargic mice show pituitary-adrenal hypercorticism. Homozygotes are smaller and weaker than their normal littermates and often die before 2 months old. Survivors of both sexes may breed, but their reproductivity is low.
| Control | ||
|---|---|---|
| Untyped from the colony | ||
| Considerations for Choosing Controls | ||
Strains carrying A allele
003301 (C57BL/6J x C3H-Eya1bor)F1/J 002083 B6 x B6EiC3 a/A-T(7;16)235Dn/J 000507 B6 x B6EiC3 a/A-Otcspf/J 000628 B6.CE-A Amy1b Amy2b/J 004200 B6;CBACa Aw-J/A-Npr2cn-2J/J 000604 B6C3 a/A-T(10;13)199H +/+ Lystbg-J/J or Lystbg-2J/J 001752 B6CBCa Aw-J/A-T(7;15)9H/J 006450 B6EiC3 a/A-Vss/J 000557 B6EiC3-+ a/LnpUl A/J 000553 B6EiC3Sn a/A-Egfrwa2 Wnt3avt/J 001811 B6EiC3Sn a/A-Otcspf-ash/J 002343 B6EiC3Sn a/A-Otcspf/J 001923 B6EiC3Sn a/A-Ts(417)2Lws Tim/J 000200 C3FeB6 A/Aw-J-Ankank/J 000638 C3FeB6 A/Aw-J-Spnb4qv-J/J 000283 LT.CAST-A/J 001759 STOCK A Tyrc Sha/J View Strains carrying A (17 strains)
Strains carrying other alleles of Cacnb4
004951 C3HeB/FeJ-Cacnb4lh-3J/J View Strains carrying other alleles of Cacnb4 (1 strain)
Strains carrying other alleles of a
View Strains carrying other alleles of a (156 strains)
Mammalian Phenotype Terms assigned by genotype
Cacnb4lh/Cacnb4lh
B6EiC3Sn a/A-Cacnb4lh/J
- immune system phenotype
- abnormal CD4-positive T cell physiology (MGI Ref ID J:115345)
- initial peak and plateau calcium responses are attenuated compared to in wild-type mice
- abnormal Peyer's patch germinal center morphology (MGI Ref ID J:5281)
- absent
- abnormal interleukin secretion (MGI Ref ID J:115345)
- IL-4 production is reduced
- decreased interleukin-2 secretion (MGI Ref ID J:115345)
- TCR-mediated IL-2 production is partially inhibited
- abnormal lymph node morphology (MGI Ref ID J:5281)
- between 18 and 35 days of age, mice exhibit fewer lymphatic follicles compared to wild-type mice
- enlarged lymph nodes (MGI Ref ID J:5281)
- mice older than 5 months exhibit enlarged lymph nodes compared to wild-type mice
- abnormal thymus development (MGI Ref ID J:5281)
- mice undergo thymus involution
- abnormal thymus lobule morphology (MGI Ref ID J:5281)
- the boundary between the cortex and medulla is not visible as in wild-type mice
- decreased Peyer's patch number (MGI Ref ID J:5281)
- decreased interferon-gamma secretion (MGI Ref ID J:115345)
- decreased lymphocyte cell number (MGI Ref ID J:5281)
- in the thymus and spleen
- decreased double-positive T cell number (MGI Ref ID J:115345)
- in mice older than 2 weeks of age when the onset of neuropathy occurs
- however, prior to 2 weeks of age mice exhibit normal T cell development
- small Peyer's patches (MGI Ref ID J:5281)
- small spleen (MGI Ref ID J:5281)
- between 15 days and 10 months of age
- small thymus (MGI Ref ID J:5281)
- between 26 and 40 days of age
- growth/size phenotype
- cachexia (MGI Ref ID J:5281)
- between 26 and 40 days of age
- digestive/alimentary phenotype
- diarrhea (MGI Ref ID J:5281)
- between 26 and 40 days of age
- vision/eye phenotype
- abnormal eye electrophysiology (MGI Ref ID J:136113)
- electroretinograms generated by retinal pigment epithelial cells are reduced compared to in wild-type mice
- maximum amplitude of the light peak is reduced and overall sensitivity is decreased compared to in wild-type mice
- hematopoietic system phenotype
- abnormal thymus development (MGI Ref ID J:5281)
- mice undergo thymus involution
- abnormal thymus lobule morphology (MGI Ref ID J:5281)
- the boundary between the cortex and medulla is not visible as in wild-type mice
- decreased lymphocyte cell number (MGI Ref ID J:5281)
- in the thymus and spleen
- decreased double-positive T cell number (MGI Ref ID J:115345)
- in mice older than 2 weeks of age when the onset of neuropathy occurs
- however, prior to 2 weeks of age mice exhibit normal T cell development
- small spleen (MGI Ref ID J:5281)
- between 15 days and 10 months of age
- small thymus (MGI Ref ID J:5281)
- between 26 and 40 days of age
The following phenotype information may relate to a genetic background differing from this JAX® Mice strain.
Cacnb4lh/Cacnb4lh
involves: BALB/cGn * C3H/HeSnJ * C57BL/6JEi
- nervous system phenotype
- *normal* nervous system phenotype (MGI Ref ID J:121463)
- synaptic electrophysiology including neurotransmitter release and end plate potentials, as well as responses to channel inhibitors is not different from wild-type NMJs
- abnormal CNS synaptic transmission (MGI Ref ID J:106959)
- peak current densities of low voltage-activated (LVA) calcium channels in thalamocortical neurons at membrane potential of -50 mV are increased by 51% compared to control
- peak current densities of high voltage calcium channels (HVA) in TC neurons are similar to wild-type
- abnormal brain wave pattern (MGI Ref ID J:79139)
- EEG of saline-treated mice display frequent polyspike discharges
- mice exhibit bilaterally synchronous electrographic bursts (seizures) that last 1.5 s with a frequency of 127 per hour
- abnormal motor neuron morphology (MGI Ref ID J:121463)
- synaptic electrophysiology including neurotransmitter release and end plate potentials, as well as responses to channel inhibitors is not different from wild-type NMJs
- abnormal neuromuscular synapse morphology (MGI Ref ID J:121463)
- neuromuscular junction area (area staining for acetylcholine receptors) in ~38% smaller relative to wild-type mice (277 um2 vs 446 um2)
- seizures (MGI Ref ID J:1484)
- mice exhibit absence seizures
- seizures are increased by treatment with GAB agonists and decreased by treatment with GABA antagonists
- behavior/neurological phenotype
- abnormal motor capabilities/coordination/movement (MGI Ref ID J:79139)
- transient periods of severe motor disability are superimposed on the more minor abnormalities
- abnormal posture (MGI Ref ID J:79139)
- mice show periods of abnormal flexion or extension of the trunk most apparent at 4 weeks of age
- ataxia (MGI Ref ID J:79139)
- mice show wide stance, imprecise limb placement, poor timing and interlimb coordination, and a staggering gait, all characteristic of ataxia
- circling (MGI Ref ID J:79139)
- mice display circling, becoming less frequent by 5-6 weeks of age
- hypoactivity (MGI Ref ID J:79139)
- mice show transient periods of reduced locomotor activity when tested in photocell chambers
- impaired coordination (MGI Ref ID J:79139)
- mice perform very poorly in rotarod tests, falling off the rod whereas wild-type remain on for ~length of test
- impaired righting response (MGI Ref ID J:79139)
- mice tend to fall and are unable to regain upright position, becoming less frequent by 5-6 weeks of age
- seizures (MGI Ref ID J:1484)
- mice exhibit absence seizures
- seizures are increased by treatment with GAB agonists and decreased by treatment with GABA antagonists
- clonic seizures (MGI Ref ID J:79139)
- mice have attacks with clonic movements of the limbs most apparent at 4 weeks of age
- tonic seizures (MGI Ref ID J:79139)
- mice show periods of tonic extension or retraction of the limbs at 4 weeks of age
- muscle phenotype
- muscle spasms (MGI Ref ID J:79139)
- mice display facial twitching, becoming less prominent by 5-6 weeks of age
- hearing/vestibular/ear phenotype
- circling (MGI Ref ID J:79139)
- mice display circling, becoming less frequent by 5-6 weeks of age
- homeostasis/metabolism phenotype
- decreased sensitivity to xenobiotics (MGI Ref ID J:1484)
- mice are insensitive to phenytoin and carbamazepine
Cacnb4lh/Cacnb4lh
BALB/cGn-Cacnb4lh
- life span-post-weaning/aging
- premature death (MGI Ref ID J:12166)
- mice exhibit high mortality rates between 3 and 4 weeks of age
- reproductive system phenotype
- absent corpus luteum (MGI Ref ID J:12166)
- reduced fertility (MGI Ref ID J:12166)
- mice exhibit reduced reproductivity
- immune system phenotype
- abnormal thymus development (MGI Ref ID J:12166)
- mice undergo thymus involution
- increased length of allograft survival (MGI Ref ID J:5766)
- between 15 to 30 days of age, mice fail to reject skin grafts as wild-type mice do
- however, by 45 days of age graft rejection is normal
- nervous system phenotype
- pituitary gland hyperplasia (MGI Ref ID J:12166)
- pituitary glands exhibit an increase in cellular population density compared to in wild-type pituitaries
- behavior/neurological phenotype
- abnormal behavior (MGI Ref ID J:28465)
- mice sit up with front feet contracted for several minutes
- abnormal fear/anxiety-related behavior (MGI Ref ID J:28465)
- mice appear nervous
- abnormal locomotor activation (MGI Ref ID J:28465)
- mice exhibit difficulty in moving forward and make long pauses between movements, often hunch the back and raise first the front foot then the hind foot
- endocrine/exocrine gland phenotype
- absent corpus luteum (MGI Ref ID J:12166)
- pituitary gland hyperplasia (MGI Ref ID J:12166)
- pituitary glands exhibit an increase in cellular population density compared to in wild-type pituitaries
- growth/size phenotype
- postnatal growth retardation (MGI Ref ID J:12166)
- hematopoietic system phenotype
- abnormal thymus development (MGI Ref ID J:12166)
- mice undergo thymus involution
Cacnb4lh/Cacnb4lh
involves: BALB/cGn
- behavior/neurological phenotype
- abnormal gait (MGI Ref ID J:28466)
- at 14 to 16 days of age, mice exhibit a mild gait instability
- convulsive seizures (MGI Ref ID J:28466)
- mice exhibit clonic and/or tonic seizures
- excitement during seizure worsens the severity of the seizure
- however, gentle handling quells the seizures until released
- sometimes mice exhibit a Jacksonian march during seizures
- lethargy (MGI Ref ID J:28466)
- when removed from a swim test mice exhibit slowed responses with hindlimbs appearing weak and he animal unable to ascend a 45 degree incline for 10 to 15 minutes
- nervous system phenotype
- abnormal nerve conduction (MGI Ref ID J:6556)
- mice exhibit adult motor nerve conduction velocities at 3 months of again instead of 5 as in wild-type mice
- reduced nerve conduction velocity (MGI Ref ID J:6556)
- peripheral motor nerves exhibit reduced conduction velocity and prolonged distal latency
- convulsive seizures (MGI Ref ID J:28466)
- mice exhibit clonic and/or tonic seizures
- excitement during seizure worsens the severity of the seizure
- however, gentle handling quells the seizures until released
- sometimes mice exhibit a Jacksonian march during seizures
- immune system phenotype
- increased IgG1 level (MGI Ref ID J:5883)
Research Applications
This mouse can be used to support research in many areas including:Cacnb4lh related
Cell Biology Research
Channel and Transporter Defects (calcium)
Endocrine Deficiency Research
Adrenal Cortex Defects
Hypothalamus/Pituitary Defects
Immunology and Inflammation Research
Immunodeficiency Associated with Other Defects
Internal/Organ Research
Adrenal Cortex Defects
Neurobiology Research
Ataxia (Movement) Defects
Channel and Transporter Defects (calcium)
Epilepsy
| Allele Symbol | A | ||
|---|---|---|---|
| Allele Name | wild type | ||
| Gene Symbol and Name | a, nonagouti | ||
| Chromosome | 2 | ||
| Gene Common Name(s) | AGSW; AGTI; AGTIL; ASP; As; MGC126092; MGC126093; SHEP9; agouti; agouti signal protein; agouti suppressor; | ||
| Allele Symbol | Cacnb4lh | ||
| Allele Name | lethargic | ||
| Common Name(s) | beta4lh; lh; | ||
| Strain of Origin | BALB/cGn | ||
| Gene Symbol and Name | Cacnb4, calcium channel, voltage-dependent, beta 4 subunit | ||
| Chromosome | 2 | ||
| Gene Common Name(s) | 3110038O15Rik; CAB4; CACNLB4; Cchb4; EA5; EJM; RIKEN cDNA 3110038O15 gene; calcium channel, B4 subunit; lethargic; lh; | ||
| Molecular Note | A four bp insertion that disrupts the consensus 5' splice site of intron A/B was identified. Reduced levels of transcripts are produced from this allele and are present at approximately 20% of the normal levels in homozygous mice. [MGI Ref ID J:38214] | ||
This strain will not have a genotyping protocol or one is not currently available.
Helpful Links
Optimizing PCR Protocols
Burgess DL; Jones JM; Meisler MH; Noebels JL. 1997. Mutation of the Ca2+ channel beta subunit gene Cchb4 is associated with ataxia and seizures in the lethargic (lh) mouse. Cell 88(3):385-92. [PubMed: 9039265] [MGI Ref ID J:38214]
Dung HC; Swigart RH. 1972. Histo-pathologic observations of the nervous and lymphoid tissues of lethargic mutant mice. Tex Rep Biol Med 30(1):23-39. [PubMed: 5037658] [MGI Ref ID J:5281]
Lin FH; Cao Z; Hosford DA. 1993. Increased number of GABAB receptors in the lethargic (lh/lh) mouse model of absence epilepsy. Brain Res 608(1):101-6. [PubMed: 8388308] [MGI Ref ID J:4388]
A related
Czyzyk TA; Sikorski MA; Yang L; McKnight GS. 2008. Disruption of the RIIbeta subunit of PKA reverses the obesity syndrome of Agouti lethal yellow mice. Proc Natl Acad Sci U S A 105(1):276-81. [PubMed: 18172198] [MGI Ref ID J:131039]Cacnb4lh relatedJackson IJ; Budd PS; Keighren M; McKie L. 2007. Humanized MC1R transgenic mice reveal human specific receptor function. Hum Mol Genet 16(19):2341-8. [PubMed: 17652101] [MGI Ref ID J:129904]
Aizawa M; Ito Y; Fukuda H. 1997. Pharmacological profiles of generalized absence seizures in lethargic, stargazer and gamma-hydroxybutyrate-treated model mice. Neurosci Res 29(1):17-25. [PubMed: 9293489] [MGI Ref ID J:43226]Badou A; Basavappa S; Desai R; Peng YQ; Matza D; Mehal WZ; Kaczmarek LK; Boulpaep EL; Flavell RA. 2005. Requirement of voltage-gated calcium channel beta4 subunit for T lymphocyte functions. Science 307(5706):117-21. [PubMed: 15637280] [MGI Ref ID J:95687]
Badou A; Jha MK; Matza D; Mehal WZ; Freichel M; Flockerzi V; Flavell RA. 2006. Critical role for the beta regulatory subunits of Cav channels in T lymphocyte function. Proc Natl Acad Sci U S A 103(42):15529-34. [PubMed: 17028169] [MGI Ref ID J:115345]
Ball SL; Powers PA; Shin HS; Morgans CW; Peachey NS; Gregg RG. 2002. Role of the beta(2) subunit of voltage-dependent calcium channels in the retinal outer plexiform layer. Invest Ophthalmol Vis Sci 43(5):1595-603. [PubMed: 11980879] [MGI Ref ID J:80080]
Burgess DL; Biddlecome GH; McDonough SI; Diaz ME; Zilinski CA; Bean BP; Campbell KP; Noebels JL. 1999. beta subunit reshuffling modifies N- and P/Q-type Ca2+ channel subunit compositions in lethargic mouse brain. Mol Cell Neurosci 13(4):293-311. [PubMed: 10328888] [MGI Ref ID J:57712]
Caddick SJ; Wang C; Fletcher CF; Jenkins NA; Copeland NG; Hosford DA. 1999. Excitatory but not inhibitory synaptic transmission is reduced in lethargic (Cacnb4(lh)) and tottering (Cacna1atg) mouse thalami. J Neurophysiol 81(5):2066-74. [PubMed: 10322048] [MGI Ref ID J:56225]
Devanagondi R; Egami K; LeDoux MS; Hess EJ; Jinnah HA. 2007. Neuroanatomical substrates for paroxysmal dyskinesia in lethargic mice. Neurobiol Dis 27(3):249-57. [PubMed: 17561408] [MGI Ref ID J:134825]
Dickie MM. 1964. Lethargic (lh) Mouse News Lett 30:31. [MGI Ref ID J:28465]
Dung HC. 1981. 2. Lethargic mice. In: Immunologic Defects in Laboratory Animals. Plenum Press. [MGI Ref ID J:28475]
Dung HC. 1977. Deficiency in the thymus-dependent immunity in lethargic mutant mice. Transplantation 23(1):39-43. [PubMed: 13525] [MGI Ref ID J:5766]
Dung HC. 1975. Growth retardation, high mortality, and low reproductivity of neurological mutant mice. Anat Rec 181:347-348 (Abstr.). [MGI Ref ID J:12166]
Dung HC; Lawson RL; Stevens M. 1977. A study of the increased serum level of IgG1 in 'lethargic' mice combined with a depressed thymus-dependent lymphoid system. J Immunogenet 4(4):287-93. [PubMed: 410895] [MGI Ref ID J:5883]
Dung HC; Swigart RH. 1972. Histo-pathologic observations of the nervous and lymphoid tissues of lethargic mutant mice. Tex Rep Biol Med 30(1):23-39. [PubMed: 5037658] [MGI Ref ID J:5281]
Herring JM; Dung HC; Yoo JH; Yu J. 1981. Chronological studies of peripheral motor nerve conduction in lethargic mice. Electromyogr Clin Neurophysiol 21(2-3):121-34. [PubMed: 7261997] [MGI Ref ID J:6556]
Hosford DA; Clark S; Cao Z; Wilson WA Jr; Lin FH; Morrisett RA; Huin A. 1992. The role of GABAB receptor activation in absence seizures of lethargic (lh/lh) mice. Science 257(5068):398-401. [PubMed: 1321503] [MGI Ref ID J:1484]
Hosford DA; Lin FH; Kraemer DL; Cao Z; Wang Y; Wilson JT Jr. 1995. Neural network of structures in which GABAB receptors regulate absence seizures in the lethargic (lh/lh) mouse model. J Neurosci 15(11):7367-76. [PubMed: 7472490] [MGI Ref ID J:29786]
Hosford DA; Wang Y; Liu CC; Snead OC 3rd. 1995. Characterization of the antiabsence effects of SCH 50911, a GABA-B receptor antagonist, in the lethargic mouse, gamma-hydroxybutyrate, and pentylenetetrazole models. J Pharmacol Exp Ther 274(3):1399-403. [PubMed: 7562514] [MGI Ref ID J:30226]
Ishige K; Ito Y; Fukuda H. 1999. Characterization of absence seizure-dependent cyclic AMP responsive element-and activator protein 1 DNA-binding activities in lethargic (lh/lh) mice. Neurosci Lett 262(1):53-6. [PubMed: 10076871] [MGI Ref ID J:56009]
Kaja S; Todorov B; van de Ven RC; Ferrari MD; Frants RR; van den Maagdenberg AM; Plomp JJ. 2007. Redundancy of Cav2.1 channel accessory subunits in transmitter release at the mouse neuromuscular junction. Brain Res 1143:92-101. [PubMed: 17320843] [MGI Ref ID J:121463]
Khan Z; Jinnah HA. 2002. Paroxysmal dyskinesias in the lethargic mouse mutant. J Neurosci 22(18):8193-200. [PubMed: 12223573] [MGI Ref ID J:79139]
Lin F; Barun S; Lutz CM; Wang Y; Hosford DA. 1999. Decreased (45)Ca(2)(+) uptake in P/Q-type calcium channels in homozygous lethargic (Cacnb4lh) mice is associated with increased beta3 and decreased beta4 calcium channel subunit mRNA expression. Brain Res Mol Brain Res 71(1):1-10. [PubMed: 10407181] [MGI Ref ID J:56507]
Lin F; Wang Y; Hosford DA. 1999. Age-related relationship between mRNA expression of GABA(B) receptors and calcium channel beta4 subunits in cacnb4lh mice. Brain Res Mol Brain Res 71(1):131-5. [PubMed: 10407196] [MGI Ref ID J:109316]
Lin FH; Cao Z; Hosford DA. 1993. Increased number of GABAB receptors in the lethargic (lh/lh) mouse model of absence epilepsy. Brain Res 608(1):101-6. [PubMed: 8388308] [MGI Ref ID J:4388]
Lin FH; Lin S; Wang Y; Hosford DA. 1999. Glutamate decarboxylase isoforms in thalamic nuclei in lethargic mouse model of absence seizures. Brain Res Mol Brain Res 71(1):127-30. [PubMed: 10407195] [MGI Ref ID J:109317]
Lin FH; Wang Y; Lin S; Cao Z; Hosford DA. 1995. GABAB receptor-mediated effects in synaptosomes of lethargic (lh/lh) mice. J Neurochem 65(5):2087-95. [PubMed: 7595494] [MGI Ref ID J:29347]
Marmorstein LY; Wu J; McLaughlin P; Yocom J; Karl MO; Neussert R; Wimmers S; Stanton JB; Gregg RG; Strauss O; Peachey NS; Marmorstein AD. 2006. The light peak of the electroretinogram is dependent on voltage-gated calcium channels and antagonized by bestrophin (best-1). J Gen Physiol 127(5):577-89. [PubMed: 16636205] [MGI Ref ID J:136113]
McEnery MW; Copeland TD; Vance CL. 1998. Altered expression and assembly of N-type calcium channel alpha1B and beta subunits in epileptic lethargic (lh/lh) mouse. J Biol Chem 273(34):21435-8. [PubMed: 9705268] [MGI Ref ID J:49244]
Qian J; Noebels JL. 2000. Presynaptic Ca(2+) influx at a mouse central synapse with Ca(2+) channel subunit mutations. J Neurosci 20(1):163-70. [PubMed: 10627593] [MGI Ref ID J:120572]
Sidman RL; Green MC; Appel SH. 1965. Lethargic, lh, recessive. In: Catalog of the Neurological Mutants of the Mouse. Harvard University Press. [MGI Ref ID J:28466]
Song I; Kim D; Choi S; Sun M; Kim Y; Shin HS. 2004. Role of the alpha1G T-type calcium channel in spontaneous absence seizures in mutant mice. J Neurosci 24(22):5249-57. [PubMed: 15175395] [MGI Ref ID J:96913]
Zhang Y; Mori M; Burgess DL; Noebels JL. 2002. Mutations in high-voltage-activated calcium channel genes stimulate low-voltage-activated currents in mouse thalamic relay neurons. J Neurosci 22(15):6362-71. [PubMed: 12151514] [MGI Ref ID J:106959]
Currently there no information available for this strain. This may be due to the supply level of this strain.
*Price(s) in US dollars ($)
Weeks of Age Price* Gender Cryorecovery Fee $1900.00
| Standard Supply | Repository-Cryopreserved. Must Be Recovered. Please refer to pricing and supply notes for further information. |
|---|---|
| Supply Notes |
|
*Price(s) in US dollars ($)
Weeks of Age Price* Gender Cryorecovery Fee $2470.00
| Standard Supply | Repository-Cryopreserved. Must Be Recovered. Please refer to pricing and supply notes for further information. |
|---|---|
| Supply Notes |
|
| Control | ||
|---|---|---|
| Untyped from the colony | ||
| Considerations for Choosing Controls | ||
| USA, Canada and Mexico - Control Pricing Information for Genetically Engineered Mutant Strains. | ||
| International - Control Pricing Information for Genetically Engineered Mutant Strains. | ||
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